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Nuclear patterns of phosphatidylinositol 4,5-and 3,4-bisphosphate revealed by super-resolution microscopy differ between the consecutive stages of RNA polymerase II transcription

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    SYSNO ASEP0601091
    Document TypeJ - Journal Article
    R&D Document TypeJournal Article
    Subsidiary JČlánek ve WOS
    TitleNuclear patterns of phosphatidylinositol 4,5-and 3,4-bisphosphate revealed by super-resolution microscopy differ between the consecutive stages of RNA polymerase II transcription
    Author(s) Hoboth, Peter (UMG-J) ORCID
    Sztacho, Martin (UMG-J) ORCID
    Hozák, Pavel (UMG-J) RID, ORCID
    Number of authors3
    Source TitleFEBS Journal - ISSN 1742-464X
    Roč. 291, č. 19 (2024), s. 4240-4264
    Number of pages25 s.
    Languageeng - English
    CountryGB - United Kingdom
    Keywordsswiss 3t3-cells ; phospholipase-c ; ctd ; phosphorylation ; speckles ; phosphoinositides ; kinase ; cells ; organization ; association ; gene expression ; nuclear architecture ; nuclear speckles ; nucleoplasm ; quantitative direct stochastic optical reconstruction microscopy dSTORM
    OECD categoryCell biology
    R&D ProjectsLM2023050 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    EF18_046/0016045 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    EF16_013/0001775 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    LX22NPO5102 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    LTC19048 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    LTC20024 GA MŠMT - Ministry of Education, Youth and Sports (MEYS)
    Method of publishingOpen access
    Institutional supportUMG-J - RVO:68378050
    UT WOS001219071800001
    EID SCOPUS85193002640
    DOI https://doi.org/10.1111/febs.17136
    AnnotationPhosphatidylinositol phosphates are powerful signaling molecules that orchestrate signaling and direct membrane trafficking in the cytosol. Interestingly, phosphatidylinositol phosphates also localize within the membrane-less compartments of the cell nucleus, where they participate in the regulation of gene expression. Nevertheless, current models of gene expression, which include condensates of proteins and nucleic acids, do not include nuclear phosphatidylinositol phosphates. This gap is partly a result of the missing detailed analysis of the subnuclear distribution of phosphatidylinositol phosphates and their relationships with gene expression. Here, we used quantitative dual-color direct stochastic optical reconstruction microscopy to analyze the nanoscale co-patterning between RNA polymerase II transcription initiation and elongation markers with respect to phosphatidylinositol 4,5- or 3,4-bisphosphate in the nucleoplasm and nuclear speckles and compared it with randomized data and cells with inhibited transcription. We found specific co-patterning of the transcription initiation marker P-S5 with phosphatidylinositol 4,5-bisphosphate in the nucleoplasm and with phosphatidylinositol 3,4-bisphosphate at the periphery of nuclear speckles. We showed the specific accumulation of the transcription elongation marker PS-2 and of nascent RNA in the proximity of phosphatidylinositol 3,4-bisphosphate associated with nuclear speckles. Taken together, this shows that the distinct spatial associations between the consecutive stages of RNA polymerase II transcription and nuclear phosphatidylinositol phosphates exhibit specificity within the gene expression compartments. Thus, in analogy to the cellular membranes, where phospholipid composition orchestrates signaling pathways and directs membrane trafficking, we propose a model in which the phospholipid identity of gene expression compartments orchestrates RNA polymerase II transcription.
    WorkplaceInstitute of Molecular Genetics
    ContactNikol Škňouřilová, nikol.sknourilova@img.cas.cz, Tel.: 241 063 217
    Year of Publishing2025
    Electronic addresshttps://febs.onlinelibrary.wiley.com/doi/10.1111/febs.17136
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